1To whom correspondence should be addressed: e-mail:
2Graduate School of Systems Life Sciences, Kyushu University,Fukuoka 812-8581, Japan
3Department of Biological Sciences, Faculty of Sciences 33, Kyushu University,Fukuoka 812-8581, Japan
4Core Research for Evolutional Science and Technology (CREST), Japan Science and Technology Agency (JST), Kawaguchi Center Building, 4-1-8 Hon-cho, Kawaguchi, Saitama 332-0012, Japan
5Department of Physiology, Tokyo Women's Medical University School of Medicine,Tokyo 162-8666, Japan
6Glycomedicine Technology Research Center (GTRC), National Institute of Advanced Industrial Science and Technology (AIST), Tsukuba 305-8568, Japan
7Present address: Department of Pharmacology, Graduate School of Medicine, Osaka University, Osaka 565-0871, Japan.
8Present address: Saitama University Brain Science Institute, Saitama 338-8570, Japan.